Sec Form 4 Filing - Walker Matthew Allen @ GreenLight Biosciences Holdings, PBC - 2023-07-24

FORM 4 Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940

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1. Name and Address of Reporting Person Walker Matthew Allen 2. Issuer Name and Ticker or Trading Symbol GreenLight Biosciences Holdings, PBC [ GRNA] 5. Relationship of Reporting Person(s) to Issuer (Check all applicable) X __ Director X __ 10% Owner _____ Officer (give title below) _____ Other (specify below) (Last) (First) (Middle) C/O GREENLIGHT BIOSCIENCES HOLDINGS, PBC, 200 BOSTON AVENUE, SUITE 3100 3. Date of Earliest Transaction (MM/DD/YY) 07/24/2023 (Street) MEDFORD, MA02155 4. If Amendment, Date Original Filed (MM/DD/YY) 6. Individual or Joint/Group Filing (Check Applicable Line) _____ Form filed by One Reporting Person __ X __ Form filed by More than One Reporting Person (City) (State) (Zip)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned 1.Title of Security (Instr. 3) 2. Transaction Date (MM/DD/YY) 2A. Deemed Execution Date, if any (MM/DD/YY) 3. Transaction Code (Instr. 8) 4. Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5) 5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4) 6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4) Code V Amount (A) or (D) Price Common Stock 07/24/2023 D 63,775 ( 1 ) D 0 D Common Stock 07/24/2023 J 22,348,123 ( 2 ) D 0 I By Builders Vision, LLC ( 3 ) ( 4 )

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned ( e.g. , puts, calls, warrants, options, convertible securities) 1. Title of Derivative Security (Instr. 3) 2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (MM/DD/YY) 3A. Deemed Execution Date, if any (MM/DD/YY) 4. Transaction Code (Instr. 8) 5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4, and 5) 6. Date Exercisable and Expiration Date (MM/DD/YY) 7. Title and Amount of Underlying Securities (Instr. 3 and 4) 8. Price of Derivative Security (Instr. 5) 9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4) Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares Stock Option (Right to Buy) $ 7.96 07/24/2023 D 65,000 ( 1 ) ( 5 ) 05/30/2032 Common Stock 65,000 $ 0 0 D

Reporting Owners Reporting Owner Name / Address Relationships Director 10% Owner Officer Other Walker Matthew Allen C/O GREENLIGHT BIOSCIENCES HOLDINGS, PBC 200 BOSTON AVENUE, SUITE 3100 MEDFORD, MA02155 X X Builders Vision, LLC 110 N.W. 2ND STREET BENTONVILLE, AR72172 X Signatures /s/ Matthew Allen Walker 07/24/2023 Signature of Reporting Person Date /s/ Lisa J. Forbes, as authorized officer of Builders Vision, LLC 07/24/2023 Signature of Reporting Person Date

Explanation of Responses:

( 1 )Disposed of in connection with the merger of Issuer with a wholly-owned subsidiary of the acquiring company, pursuant to which the Issuer became a wholly-owned subsidiary of the acquiring company and all outstanding shares of Issuer's Common Stock were automatically cancelled in exchange for cash payment of $0.30 per share. ( 2 )Disposed of pursuant to a contribution and exchange agreement entered into in connection with the merger described above in exchange for the same number of shares of the acquiring company with a value of $0.40 per share. ( 3 )Total number of shares beneficially owned by the Reporting Person includes: (i) 11,551,245 shares held by S2G Builders Food & Agriculture Fund III, LP ("Fund III"); (ii) 2,087,043 shares held by S2G Ventures Fund I, L.P. ("Fund I"); (iii) 8,582,284 shares held by S2G Ventures Fund II, L.P. ("Fund II" and, together with Fund I, and Fund III, the "S2G Funds"); and (iv) 127,551 shares held by Builders GRNA Holdings, LLC ("SPV"). Builders Vision, LLC is the Manager Fund I, Fund II, and SPV and the General Partner of Fund III, and has power to vote or direct the voting of shares held by the S2G Funds and SPV. The General Partners of Fund I and Fund II are S2G Ventures, LLC and S2G Ventures II, LLC, respectively. ( 4 )(Continued) Mr. Walker, a director of the Issuer, is a Managing Director of Builders Vision, LLC, the impact platform founded by Lukas T. Walton, which includes S2G Ventures. By virtue of the foregoing, S2G Ventures, LLC and S2G Ventures II, LLC, Mr. Walker and Mr. Walton may be deemed to indirectly beneficially own (as defined in Rule 13d-3 of the Exchange Act) the shares of Common Stock held by the S2G Funds. Mr. Walker and Mr. Walton each disclaims beneficial ownership of these shares of Common Stock except to the extent of any pecuniary interest therein. ( 5 )25,000 shares underlying the option vested or shall vest in four equal quarterly installments, the first of which shall vest on May 3, 2022. The remaining 40,000 shares underlying the option vested or shall vest in twelve equal quarterly installments, the first of which shall vest on May 3, 2022. The option was canceled pursuant to the merger for no payment.

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.

* If the form is filed by more than one reporting person, see Instruction 4(b)(v).

** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).

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