Sec Form 4 Filing - Spectrum Equity Investors V L P @ Ancestry.com Inc. - 2012-12-28

FORM 4 Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

UNITED STATES SECURITIES AND EXCHANGE COMMISSION Washington, D.C. 20549 STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940

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1. Name and Address of Reporting Person Spectrum Equity Investors V L P 2. Issuer Name and Ticker or Trading Symbol Ancestry.com Inc. [ ACOM] 5. Relationship of Reporting Person(s) to Issuer (Check all applicable) _____ Director X __ 10% Owner _____ Officer (give title below) _____ Other (specify below) (Last) (First) (Middle) 333 MIDDLEFIELD ROAD, SUITE 200 3. Date of Earliest Transaction (MM/DD/YY) 12/28/2012 (Street) MENLO PARK, CA94025 4. If Amendment, Date Original Filed (MM/DD/YY) 6. Individual or Joint/Group Filing (Check Applicable Line) _____ Form filed by One Reporting Person __ X __ Form filed by More than One Reporting Person (City) (State) (Zip)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned 1.Title of Security (Instr. 3) 2. Transaction Date (MM/DD/YY) 2A. Deemed Execution Date, if any (MM/DD/YY) 3. Transaction Code (Instr. 8) 4. Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5) 5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4) 6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4) Code V Amount (A) or (D) Price Common Stock, $0.001 par value 12/28/2012 D 13,263,819 ( 1 ) ( 2 ) ( 3 ) ( 4 ) ( 5 ) ( 6 ) D $ 32 0 D

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned ( e.g. , puts, calls, warrants, options, convertible securities) 1. Title of Derivative Security (Instr. 3) 2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (MM/DD/YY) 3A. Deemed Execution Date, if any (MM/DD/YY) 4. Transaction Code (Instr. 8) 5. Number of Derivative Securities Acquired (A) or Disposed of (D) (Instr. 3, 4, and 5) 6. Date Exercisable and Expiration Date (MM/DD/YY) 7. Title and Amount of Underlying Securities (Instr. 3 and 4) 8. Price of Derivative Security (Instr. 5) 9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4) 10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4) 11. Nature of Indirect Beneficial Ownership (Instr. 4) Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares

Reporting Owners Reporting Owner Name / Address Relationships Director 10% Owner Officer Other Spectrum Equity Investors V L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X SEA V Management LLC 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X SEI III ENTREPRENEURS FUND L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X SEI III Entrepreneurs LLC 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X Spectrum Equity Associates III L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X Spectrum Equity Associates V LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X SPECTRUM EQUITY INVESTORS III L P 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X SPECTRUM III INVESTMENT MANAGERS FUND LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X Spectrum V Investment Managers Fund LP 333 MIDDLEFIELD ROAD SUITE 200 MENLO PARK, CA94025 X Signatures /s/ William C. Stern, Attorney-in-Fact 01/02/2013 Signature of Reporting Person Date

Explanation of Responses:

( 1 )This Form 4 should be read in conjunction with the other Form 4 filed by Spectrum Equity Investors V, L.P. ("SEI V") as of the date hereof. Each of such Forms 4 reports the total number of shares of common stock sold by SEI V and its affiliates as described below. The amounts set forth in the two Forms 4 should not be added to reach a total. ( 2 )Consists of an aggregate of 13,263,819 shares disposed of in connection with the acquistion of Ancestry.com Inc. by a company controlled by investment funds advised by Permira Advisers, LLC on December 28, 2012 (the "Merger"), including 9,516,845 shares disposed of by SEI V, the general partner of which is Spectrum Equity Associates V, L.P., the general partner of which is SEA V Management, LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker and Christopher T. Mitchell exercise voting and dispositive power; 44,941 shares disposed of by Spectrum V Investment Managers' Fund, L.P. ("IMF V"), the general partner of which is SEA V Management, LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker and Christopher T. Mitchell exercise voting and dispositive power; ( 3 )and including 3,395,981 shares disposed of by Spectrum Equity Investors III, L.P. ("SEI III"), the general partner of which is Spectrum Equity Associates III, L.P., over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power; and 238,252 shares disposed of by SEI III Entrepreneurs' Fund, L.P. ("Entrepreneurs' III"), the general partner of which is SEI III Entrepreneurs' LLC, over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power; ( 4 )and including 57,067 shares disposed of by Spectrum III Investment Managers' Fund, L.P. ("IMF III," and together with SEI V, IMF V, SEI III and Entrepreneurs' III, the "Spectrum Funds"), over which Brion B. Applegate, William P. Collatos, Kevin J. Maroni and Randy J. Henderson exercise voting and dispositive power and are the general partners, and 6,606 shares disposed of by The Brion B. Applegate Trust, over which Brion B. Applegate exercises voting and dispositive power, 3,300 shares disposed of by William P. Collatos and 827 shares disposed of by Randy J. Henderson. ( 5 )Each of the controlling entities, individual general partners and managing directors of the Spectrum Funds, as the case may be, including Mr. Parker who is a managing director of the general partner of the general partner of SEI V and a managing director of the general partner of IMF V, and served on Ancestry.com Inc.'s board of directors until the effective time of the Merger, Brion B. Applegate, William P. Collatos, Kevin J. Maroni, Randy J. Henderson, Michael J. Kennealy, Victor E. Parker and Christopher T. Mitchell disclaims beneficial ownership of these shares except to the extent of any pecuniary interest therein. ( 6 )In connection with the transactions disclosed, certain of the Reporting Persons transferred certain of their shares of common stock to wholly owned subsidiaries before the disclosed transactions occurred.

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.

* If the form is filed by more than one reporting person, see Instruction 4(b)(v).

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